Associated species include Ulmus americana, Acer saccharum, Fagus grandifolia, Quercus rubra, Ostrya virginiana, Caulophyllum thalictroides, Hydrophyllum virginianum, Erythronium albidum, Carex hirtifolia, Adiantum pedatum, Geranium maculatum, Dentaria laciniatum, Sanguinaria canadensis, and Podophyllum peltatum.