Associated species include Populus tremuloides, Alnus rugosa, Cornus stolonifera, Ribes hirtellum, Salix petiolaris, S pyrifolia, Rubus idaeus ssp strigosus, R pubescens, Athyrium filix-femina, Calamagrostis canadensis, Epilobium coloratum, Impatiens capensis, Polygonum sagittatum, Equisetum sylvaticum, Dryopteris cristata, Galium asperellum, Mentha arvensis v villosa, and Stellaria longifolia.