Associated species include Acer saccharum, A rubrum, Quercus alba, Q rubra, Carya ovata, Populus tremuloides, P grandidentata, Carpinus caroliniana, Hepatica acutiloba, H americana, Sanguinaria canadensis, Lycopodium species, Brachyelytrum erectum, Adiantum pedatum, Smilax hispida, Orchis spectabilis, Carya cordiformis, Podophyllum peltatum, Tilia americana, Fagus grandidentata, Trillium species, Ulmus rubra, and Hydrophyllum species.