Associated species include Laportea canadensis, Podophyllum peltatum, Scutellaria ovata, Lithospermum latifolium, Arabis shortii, Silene virginica, Cacalia muhlenbergii, Cryptotaenia canadensis, Circaea quadrisulcata, Geranium maculatum, Hydrophyllum virginianum, Jeffersonia diphylla, Hydrastis canadensis, Dirca palustris, Arabis shortii, and Ulmus thomasii.